
Name: Lutolisk (Parasiluro reptolimus)
Creator: Cube67
Ancestor: Scarletshell Sandhopper
Habitat: Cube Tropical Watershed, Glicker-Rhino Tropical Watershed, Bot Tropical Watershed, Glicker-LadyM Tropical Watershed, Cube Swamp, Bot Swamp, Glicker Tropical Rainforest
Size: 38 cm
Support: Exoskeleton (Bone shielded by keratin)
Diet: Adults: Carnivore (Mattebelly, Fluterump, Villati, Common Namnder, Strontode); Larvae: Scavenger, Detritivore
Respiration: Active (Unidirectional lungs)
Thermoregulation: Ectothermic
Reproduction: Sexual (Oviparous, 2 sexes, Internal fertilization)
Finding themselves in a strange new environment, the sandhoppers of Glicker were forced to adapt rapidly. Within only a short amount of time, one population of them began to deviate relatively quickly from the ancestral form. The result of this, while not particularly different from its ancestor genetically, may be one of the most strange-looking organisms to date on Sagan 4: the Lutolisk.
Physical AdaptationsIn order to better serve as a proper walking lumb, the lutolisk’s frontmost pseudo-toe has hypertrophied into a long spike, allowing it to pull itself across dry land and riverbed alike. In order to facilitate this mode of locomotion, a new exoskeletal plate has evolved on the bottom of its caudal fin, forming a bony “sled”. This feature has a similar genetic basis to the small bony knob on the end of the grovestalker’s tail fin, with its curved shape serving to keep its precious reproductive-tied anal fin from scraping on the ground. Additionally, the tail has truncated significantly in order to further reduce friction with the ground. This also has the positive side effect of reducing the amount of bare skin on its body, meaning that it doesn’t dry out quite as quickly. However, the adult lutolisk’s strange body shape and rigid exoskeleton makes it a poor swimmer, so when they are fully submerged underwater they usually just crawl around on the bottom. Their torso is roughly barrel-shaped (except slightly laterally compressed), with a somewhat flatter area on the underside.
To suit their slightly more terrestrial lifestyle, lutolisks’ lung chambers have become larger and more vascularized, albeit with a smaller external plate. Though their gills may atrophy away as they age, lutolisks retain large gill chambers into adulthood, which now store additional air much like a Terran bird’s air sacs. As with the scarletshell sandhopper, the gill chambers (or, in this case, air sacs) are connected to the dorsal lung. Also like their ancestors, lutolisks retain two pairs of chamber openings nestled between the lung plate and thoracic bone. However, unlike their ancestors, all of these openings are primarily used for exhalation, while the nostrils are used for inhalation. This innovation allows lutolisks to breathe unidirectionally, much like their grovestalker kin.
Hunting and FeedingBecause of its cold-bloodedness and somewhat awkward, jerky locomotion, the lutolisk is primarily an ambush predator. Its purple-and-black-striped shell helps it blend in with stands of purpleflora, concealing it from its favored prey: terrestrial carpolantaians. When an ill-fated carpolantaian (or a small namnder, which are similarly soft-bodied) comes too close, the lutolisk quickly grabs it with its tentacles, using its sharp teeth to puncture and hold onto its prey. If the prey is too large to swallow whole, the lutolisk will use its firm grip to dismember it. While the lutolisk’s gut is slightly more specialized for its diet, it does not differ too much from its ancestor’s, mainly because their ancestors already had a diet consisting of fully and partially soft-bodied prey, including aquatic carpolantaians.
Behavior and PerceptionLutolisks are solitary, only ever spending time around each other in order to mate. While lutolisks do not have defined territories, they are usually aggressive towards other members of their species as adults, especially if they are of the same sex. If two lutolisks do encounter each other in the wild, they will spread out their tentacles and slowly wave them around, intimidating each other with their sharp teeth until one of them retreats. This is used as a warning signal in order to avoid further conflict, comparable to baboons baring their teeth, cats arching their back, or iguanas bobbing their head when threatened.
This species doesn’t really burrow anymore, but do partially conceal themselves in mud or sand when there isn’t many purpleflora to hide in (or if the only kind around is purple spheres, which it does coincidentally resemble when only its knees, eyes, and lung are sticking out).
The lutolisk has vertical slit-shaped pupils, giving it highly motion-sensitive vision. Its color vision is only mediocre, being most sensitive in the red range in order to discern its blue prey from surrounding purple flora. It is also able to quickly discern between dull and glossy surfaces, so that it does not get lost in patches of crystal flora where it is most visible. Lutolisks’ eyes are placed high up on their heads and are angled slightly forwards, allowing it to get a good look at its prey and not have its vision be obscured by its legs. Lutolisks have an excellent sense of smell, using it as their primary means of detecting prey and predators from afar. This is to compensate for their lack of ears, as the only way the lutolisk is able to ‘hear’ is by detecting vibrations in the ground using its tail bone.
Life CycleLike their ancestors, the lutolisk is still bound to the water by its reproduction, although due to improved osmoregulation they are now able to have their offspring in brackish and freshwater. They mate belly-to-belly, partially or mostly submerged in water. The female lays many small eggs into a small depression underwater and then covers them with a small pile of sand using her tail. The larvae initially look like a slightly rounder and more truncated version of their ancestors’ larvae. Like the larvae of other amphibious siluros, lutolisk larvae have a dorsal lung, but it’s small and filled with water from the gill chambers. As in their ancestors, the larvae take time to develop their exoskeleton. Only when their shell is partially grown do they start showing the signature traits of a lutolisk: their front finger starts to elongate, their tail shortens even more, and their lung plate takes on a slightly conical shape. It is during this stage that they stop swimming as much and begin to switch to a lifestyle resembling that of their adults, although they are not considered fully mature until they stop eating carrion and grow twofold from there to reach their full size. Lutolisks can live for around 20 years but usually die before they reach that age.

(Supplementary image of head)

(Sketch of isometric and front views for the gallery)
POST-SUBMISSION NOTE: this thing is supposed to camouflage within stands of Stellafrutex so maybe approve that first