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Does this overlap with Ramul?

This is not a beach species, it lives exclusively out at sea.

How does having a septum-divided butt-nostril help it breathe while flying? Does it inhale through one and out the other? It get the advantage that it can still breathe if it gets clogged for some reason, but the other part of the logic doesn't make sense.

I figure petrolignum would be heavier than its ancestor's beak, but you did mention a weight-saving feature.

This is an interesting organism. Are all the pearly parts of its body petrolignum, or just its beak?

It's petrolignin, as in lignin. Petrolignum is a genus of aquatic crystal flora in Beta.

The general thought is that the septum makes the breathing more out the side than backwards. When flying at full speed, air currents should make it take more energy to inhale with how the butt nostril is positioned in most plents.

All the pearly parts are petrolignin, yes. As stated in the description, all exposed wood is replaced. I conceptualized it as being like a basic material replacement / alteration mutation for all exposed wood surface (which is probably distinguished from the wood surface that muscles attach to in the genes coding for what materials go where).