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When I read "crafted tools", I interpreted that as not meaning prosthetic limbs or teeth, but simply wielded tools which fulfilled similar functions. Perhaps the female would use spears, perhaps small ones, to tenderize or dismember meat to replace the function of her fangs.

I've added details about it. I did intend that tools would be replacing the function, not serving as a prosthetic (though that did give me an idea, which I also added)

Isn't the section about the captive females escaping big enough to justify splitting it off from the "Rogue Reproductive Behavior" section?

It's interesting to see them using prosthetic limbs, even if it's rare behavior. I wonder if this could inspire some fictionalized account of a Twineshrog missing a limb killing or mutilating other Twineshrogs for their limbs. (I already wrote some dialogue, though who knows when I'll finish it.)

Is it realistic for them to recognize equivalent function to replace their limbs, but not be sophonts? Do they consider their own limbs tools? Do they simply think, "Oh, look, so that's where I left my tail---on a dead shrog's body" and attach it where the tail goes? Do they simply think "that's where a tail goes" as they would regard logs on a nest, or "flat shrog in water look bad, no tail, make it look right, add tail"?

Tool use is already an act of instinctively using objects as an extension of one's body. In the right situation, non-sophont animals with crafted prosthetics are not out of the question even on Earth. I figure they know it's not really their tail, but they can also intuitively know they can use it like it is.

Changes in behavior are good. Approved