What a shocking development we have going on here.

There's a space after "Thorny Toadtuga" in the diet list.

Odd..."lesser Argusraptor" really is capitalized that way on the wiki page for the Argusraptor Complex. It was the terrible Argusraptor that had odd capitalization. I'm unsure what the protocol is for named subspecies.

"Smaller prey like Toadtuga at first," It seems you forgot to add something like "They will hunt" to the beginning.

"their spike" doesn't each cheek pouch have a single spike each? Wouldn't "spikes", plural, be appropriate? Would the spike getting in the way when moving through narrow areas so significantly affect Valley Constrictors' survival and reproduction as to be lost entirely? Then again, it's just a spike, not a complex internal structure.

"To maximize genetic diversity" suggests they consciously understand genetic diversity, which is unlikely. I suggest, "they are all instinctively driven to breed in this valley, and the mixing of the populations' genetics there ensures genetic diversity."

"M and females" Either a male named M is keeping all the females to himself, or this is a typo.

I don't know about others, but I capitalize Argusraptor in descriptions regardless of other capitalization because it's the name of the genus.